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MS
unreviewed

Glucose-6-phosphatase (G6PC + G6PT translocase)

2 reactions · 2 pathways

Cofactors used

Clinical / pharmacological. von Gierke disease (GSD I): fasting hypoglycaemia, lactic acidosis, hyperuricaemia

What it does, reaction by reaction

2 reactions

Gluconeogenesis Carbohydrate Metabolism · Cytosol + mitochondrion + ER

step 13 Irreversible

Glucose-6-phosphate + H2O → glucose + Pi

Converts Glucose-6-phosphate H2O into Glucose Pi

Notes

The endoplasmic-reticulum glucose-6-phosphatase system hydrolyzes glucose-6-phosphate after transport into the ER lumen by glucose-6-phosphate translocase; the glucose and phosphate are returned to the cytosol through transporters. Hydrolysis requires water and is the final bypass of hexokinase/glucokinase, is irreversible, and permits hepatic or renal release of free glucose.

Glycogenolysis (Glycogen Breakdown) Carbohydrate Metabolism · Cytosol + ER (liver)

step 5 Irreversible

Glucose-6-phosphate + H2O → glucose + Pi

Converts Glucose-6-phosphate H2O into Glucose Pi

Notes

In liver and renal cortex, the endoplasmic-reticulum glucose-6-phosphatase system hydrolyzes glucose-6-phosphate after its transport into the ER lumen. This irreversible step enables release of glucose to blood; it is absent from skeletal muscle, explaining why muscle glycogen cannot directly correct hypoglycemia.

Showing all 2 reactions.

What accelerates and inhibits it

Regulation is pathway-specific, so each context is listed separately

3 entries

Gluconeogenesis

Listed there as: Glucose-6-phosphatase

Accelerated by

Substrate availability

Inhibited by

Hormonal control

Induced by glucagon and cortisol during fasting; repressed by insulin in the fed state

Glycogenolysis (Glycogen Breakdown)

Listed there as: Glucose-6-phosphatase (liver/kidney)

Accelerated by

Substrate availability

Inhibited by

Hormonal control

Expression rises in fasting with glucagon/cortisol and falls with insulin

Pentose Phosphate Pathway (Oxidative and Non-oxidative phases)

Listed there as: Glucose-6-phosphate dehydrogenase

Accelerated by

NADP+, glucose-6-phosphate; increased NADP+/NADPH ratio

Inhibited by

NADPH

Hormonal control

Insulin induces expression in liver and adipose tissue during lipogenesis

Recent literature

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